Shahdan, 2007: 22). North Bornean representatives of Puntioplites bulu appear more slender and lack the short dark bars on their body. They are distinguished from Puntioplites waandersii in having 7 rows of scales (versus 9) between the dorsal fin origin and lateral line, and 16 (versus 20) circumpeduncular scales (Kottelat et al., 1993: 42).
Sumber: Annotated checklist of freshwater fishes from Brunei Darussalam, Borneo
